Selected Publications:

(click on pdf to download)

Hanson, T. 2018. Biodiversity conservation and armed conflict: a warfare ecology perspective. Annals of the New York Academy of Sciences 1429: 50-65. (pdf)

Hanson, T., and J.S. Ascher. 2018. An unusually large nesting aggregation of the digger bee Anthophora bomboides Kirby, 1838 (Hymenoptera: Apidae) in the San Juan Islands, Washington State. Pan-Pacific Entomologist 94: 4-16.

Hanson, T., G. J. Wiles, and J. K. Gaydos. 2016. A novel public-private partnership model for improving the listing of endangered species. Biodiversity and Conservation 25: 193-198. (pdf)

Hanson, T., and G. J. Wiles. 2015. Washington State Status Report for the Tufted Puffin.
Washington Department of Fish and Wildlife, Olympia, WA, 62 pp. (pdf


Hanson, T., and G. Machlis. 2014. The broad view of warfare ecology: response to Marler. Environmental Conservation 41: 241. (pdf)


Hudson, L. N., T. Newbold, S. Contu, S. L. L. Hill, et al. 2014. The PREDICTS database: a global database for how terrestrial biodiversity responds to human impacts. Ecology and Evolution doi: 10.1002/ece3.1303. (pdf)

Pfeifer, M., V. Lefebvre, L. Baeten . . . T. Hanson, et al. 2014. BIOFRAG – a new database for analysing biodiversity response to forest fragmentation. Ecology and Evolution doi: 10.1002/ece3.1036. (pdf)


Potter, A., T. Hanson, and S. Vernon. 2011. Surveys for the Island Marble Butterfly (Euchloe
ausonides insulanus) in San Juan County, Washington, 2010. Washington Department of Fish and Wildlife, Olympia, WA, 23 pp.


Hanson, T. 2011. Macrolichens of the Pacific Northwest (book review). Western North
American Naturalist 71: 570-571. (pdf)

Machlis, G., Hanson, T., Spiric, Z., and J. McKendry, (eds). 2011. Warfare Ecology: A New Synthesis. Springer, Dordrecht, Netherlands.


Hanson, T., A. Potter, and S. Vernon. 2010. Surveys for the Island Marble Butterfly (Euchloe
ausonides insulanus) in San Juan County, Washington, 2009. Washington Department of Fish and Wildlife, Olympia, WA, 32 pp.


Machlis, G.E. and T. Hanson. 2009. Sobre la ecologia de los conflictos armados. Papeles 105: 11-23.


Hanson, T., Brooks, T.M., da Fonseca, G.A.B., Hoffman, M., Lamoreaux, J.F., Machlis, G., Mittermeier, C.G., Mittermeier, R.A., Pilgrim, J.D. 2009.  Warfare in biodiversity hotspots. Conservation Biology: 23: 578-587. (pdf)


Donovan, S., Looney, C., Hanson, T., Sanchez-deLeon, Y., Wulfhorst, J. D., Eigenbrode, S., Jennings, M., Johnson-Maynard, J. and Bosque-Perez, N. 2009. Linking social and biophysical  conservation perspectives in an endangered ecosystem: the Palouse as a model for bioregional planning. Ecology and Society: 14: 9. (URL: (pdf)


Hanson, T., A. Potter, J. Miskelly, and S. Vernon. 2009. Surveys for the Island Marble (Euchloe ausonides insulanus) in San Juan County, Washington, 2008. Washington Department of Fish and Wildlife, Olympia, WA, 30 pp.


Machlis, G., and T. Hanson. 2008. Warfare ecology. Bioscience 58: 729-736. (pdf)


Hanson, T., Sanchez-deLeon, Y., Johnson-Maynard, J., and Brunsfeld, S. 2008. The influence of soil and site characteristics on Palouse prairie plant communities. Western North American Naturalist 68: 231-240. (pdf)


Hanson, T.R., Brunsfeld, S.J, Finegan, B. and L.P. Waits. 2008. Pollen dispersal and genetic structure of the tropical tree Dipteryx panamensis in a fragmented landscape. Molecular Ecology 17: 2060-2073. (pdf)


Hanson, T. 2008. First observation of duetting in the Olive-backed Euphonia (Euphonia gouldi). Wilson Journal of Ornithology 120: 200-201. (pdf)


Hanson, T.R., Brunsfeld, S.J, Finegan, B. and L.P. Waits. 2008. Characterization of microsatellite primers for the almendro (Dipteryx panamensis), a tetraploid rainforest tree. Molecular Ecology Resources 8: 425-427. (pdf)


Hanson, T. 2007. Consumption of Carapa guianensis bark by Cebus capucinus in La Reserva Biologica
Indio-Maiz, Nicaragua. Neotropical Primates 14: 85-87. (pdf)


Hanson, T., Brunsfeld, S., Finegan, B. and L. Waits. 2007. Conventional and genetic measures of seed dispersal for Dipteryx panamensis (Fabaceae) in continuous and fragmented Costa Rican rainforest. Journal of Tropical Ecology 23: 635-642. (pdf)


Hanson, T., Newmark, W. and W. Stanley. 2007. Forest fragmentation and predation on artificial nests in the Usambara Mountains, Tanzania. African Journal of Ecology 45: 499-507. (pdf)


Hanson, T. 2006.  Effects of habitat fragmentation on the reproductive ecology and conservation genetics of the almendro (Dipteryx panamensis), a keystone rainforest tree.  Ph.D. Dissertation, University of Idaho, Moscow, Idaho.


Hanson, T., Brunsfeld, S., and B. Finegan. 2006. Variation in seedling density and seed predation indicators for the emergent tree Dipteryx panamensis in continuous and fragmented rainforest. Biotropica 38: 770-774. (pdf)


Hanson, T.  2006.  To predate or depredate: what’s the word? Bulletin of the Ecological Society of America 87: 128-131. (pdf)


Hanson, T. 2006. First observations of incubation behavior for the Stripe-breasted Wren (Thryothorous
thoracius). Ornitologia Neotropical 17: 453-456. (pdf)


Hanson, T. 2000. Effects of Forest Disturbance and Fragmentation on Nest Predation in the Usambara Mountains, Tanzania. M.S. Thesis, University of Vermont, Burlington, Vermont.

Thor's research interests include birds, plants, primates, insects, endangered species, and the interface between human and natural systems. He has studied the effects of habitat fragmentation on rare plant communities, avian nest predation, and the reproductive ecology and genetic diversity of rainforest trees.  Thor is currently involved in projects on Tufted Puffins and native bees, and his work on the interface between warfare and biodiversity conservation has been covered by news outlets around the world, including Reuters, Scientific American, The New York Times, The Huffington Post, and The Times of India.

Thor Hanson - Author & Biologist